7. Suppose that the populations
y
of a certain species of fish in a given
area of the ocean is described by the logistic equation
dy
dt
=
r
(1
−
y
K
)
y.
Although it is desirable to utilize this source of food, it is intuitively
clear that if too many fish are caught, then the fish population may be
reduced below a useful level and possibly even driven to extinction. The
next few problems explore some of the questions involved in formulating
a rational strategy for managing the fishery.
(a) At a given level of effort, it is reasonable to assume that the rate
at which fish are caught depends on the population
y
: the more
fish there are, the easier it is to catch them. Thus we assume that
the rate at which fish are caught is given by
Ey
, where
E
is a
positive constant, with units of 1/time, that measures the total
effort made to harvest the given species of fish.
To include this
effect, the logistic equation is modified and replaced by
dy
dt
=
r
(1
−
y
K
)
y
−
Ey.
This equation is known as the Schaefer model after the biologist
M. B. Schaefer, who applied it to fish populations.
i. Show that if
E < r
, then there are two equilibrium points,
y
1
= 0 and
y
2
=
K
(1
−
E/r
)
>
0.
ii. Show that
y
1
is unstable and
y
2
is asymptotically stable.
iii. A sustainable yield
Y
of the fishery is a rate at which fish can
be caught indefinitely.
It is the product of the effort
E
and
the asymptotically stable population
y
2
. Find
Y
as a function
of the effort
E
; the graph of this function is known as the
yield-effort curve.
iv. Determine
E
so as to maximize
Y
and thereby find the maxi-
mum sustainable yield
Y
m
.
(b) In this problem we assume that fish are caught at a constant rate
h
independent of the size of the fish population. Then
y
satisfies
dy
dx
=
r
(1
−
y/K
)
y
−
h.
The assumption of a constant catch rate
h
may be reasonable when
y
is large but becomes less so when
y
is small.
i. If
h < rK/
4, show we have two equilibrium points
y
1
and
y
2
with
y
1
< y
2
; determine these points.
ii. Show that
y
1
is unstable and
y
2
is asymptotically stable.
iii. From a plot of
f
(
y
) versus
y
, show that if the initial population
y
0
> y
1
, then
y
→
y
2
and
t
→ ∞
, but that if
y
0
< y
1
, then
y
decreases as
t
increases. increases. Note that
y
= 0 is not
an equilibrium point, so if
y
0
< y
1
, then extinction will be
reached in a finite time.
2